| Ben Stiller | Alex (hlas) | |
| David Schwimmer | Melman (hlas) | |
| Sacha Baron Cohen | King Julien XIII (hlas) | |
| Andy Richter | Mort (hlas) | |
| Bryan Cranston | Vitaly (hlas) | |
| Jessica Chastain | Gia (hlas) |
| Chris Rock | Marty (hlas) | |
| Jada Pinkett Smith | Gloria (hlas) | |
| Cedric the Entertainer | Maurice (hlas) | |
| Frances McDormand | Captain Chantel DuBois (hlas) | |
| Martin Short | Stefano (hlas) | |
| Paz Vega | Carmen (hlas) |

